Other than the folks working on the Nipigon River population the general data indicates that coaster brook trout are largely excess production the emigrates from flash basin streams in good production years where YOY fish exceed carrying capacity and end-up in open lake waters. USGS, and MiDNR researchers tagged native brook trout and planted coaster strain fish with small electromagnetic insert transmitters that generated two distinct signals in three streams that discharge into Lake Superior. They laid electromagnetic generating cables in the substrate at the mouths of these streams and recorded annual emigration rates of each brook trout stock. Over the duration of the study native brook trout emigrated to open lake waters at a higher rate seasonally and annually that the coaster strain plant fish, largely negating the genetics driven hypothesis with regard to what makes a brook trout a coaster brook trout. The more abundant food resources (many of the Lake Superior tributaries that contain basins consisting largely of Canadian Shield igneous rock formations also form anchor ice deposits in severe winter weather intervals, diminishing primary and secondary production and productivity.
Niche overlap and degree of annual resource competition among Great Lakes salmonines, stratified by open lake quadrant in Lake Michigan was quite convincingly addressed by Matt Kornis, USFWS researcher at their Green Bay Facility:
(PDF) Spatiotemporal patterns in trophic niche overlap among five salmonines in Lake Michigan, USA (researchgate.net) Those of us who argued that the MDNR, WDNR, IDNR, and Illinois.DNR managers made a massive mistake when they collectively opted to choose a fifty percent reduction in chinook plants (Option 2) over the option of applying a multispecies plant reduction approach based on bioconversion and consumption efficiency data (Option 4) in spite of each Decision Analysis Model scenario "run" one hundred, one year iterations, yielding a significantly higher probability of generating a great alewife population stock via the muli-species reductions when compared to the single species reduction option. What drove their decision? They would not have to toss fish in the hatchery rearing systems that were species dependent on multi-year rearing cycles.
They saved some hatchery fish.. avoiding having to explai to the politicians why they threw the in a landfill.and nearly crashed the entire open lake population, very nearly destroying a multi-billion dollar fishery!
Pre-dreissenid mussel invasion, spring diatom blooms largely drove the productivity of inshore Great Lakes waters providing high free fatty acid content forage for cladoceran zooplankton as well as the inshore native Gammarid amphipods and their deepwater cousin, Diporeia sp., as well as Mysis diluviana stocks. Post-spawn alewife would move offshore into waters > 150 FOW to feed on Diporeia sp. and Mysis sp. to regain somatic energy stores that they had "tapped" to make gametes Coincidentally, Diporeia sp. and Mysis sp., as well as cladoceran zooplanktors are at their annual peak in proportional free fatty acid concentrations in body tissues in mid-Ju;y. Dreissenid mussel colony filter feeding has markedly impacted bothe the scope and duration of the inshore diatom blooms in the lower Great Lakes that they infest now. In 2007 there were roughly 9 age-classes of alewife within the Lake Michigan stock. Doctor Kornis was able to document, via stable isotope food habit assessment analysis techniques that lake trout stocks that surpassed Pacific salmon stock numbers over the 2007-2009 interval feed preferentially on these adult (3YO and older fish) alewife, while most of the remaining salmonines have converted to feeding on the more numerous juveniles stock component, including chinook, the largest annual alewife consumer population in open lake waters. Consequently, driving the shift in the age structure of the alewife stock to one consisting of age-! and Age-!! fish and a smattering of adults carrying the open lake population in Lake Michigan. MSU's Quantitative Fisheries Center has now developed a multi-species predator-prey model based on species specific salmonine predator stock and prey forage stock biomass that is replacing the old single species Chinook predaotor-alewife prey model used since 2013 to inform species specific annual plant rates, as well as co-factoring in wild origin production of chinook, coho, lake trout, and steelhead.